Not all articles are created equal.  Even the best SEO focused article will fail to produce any appreciable income unless the keywords chosen fit with those that visitors are highly motivated and interested in.  For example, while an article on the topic of trombone repair might pull in good traffic, very few people may be likely to click on any advertisements in the article due to the fact that people searching for trombone repair information may not be interested in buying a new trombone or buying the new trombone accessories that are advertised via the Adsense ads or others. 

SEO, KEI, and Common Sense
Writing article content that not only draws traffic but also draws advertising clicks involves selecting keywords with high KEI scores coupled with fluid, dynamic writing that guides the keywords towards topics that users are actively looking to buying in the near future.  An excellent example of this phenomenon is an article that is written as a review of a current product with mass appeal.  Simply by its nature such an article will pull in more more potential buyers of that product.  At this point a writer can take the article in two different directions, either answer all of the potential questions a buyer might have and supply affiliate links as a primary means of revenue or provide more general information and hope that the served ads provide additional information that makes the user click them to learn more.
 
Finding a profitable balance between writing for SEO and writing what users actually want to find is a very complicated process.  As with many things the best way to find what works best for an individual web site is to monitor an article's success and then make adjustments to attempt to improve that performance.